Zoo chiefs have hailed a successful 12 months as they marked a year since two giant pandas arrived in the UK.
Tian Tian and Yang Guang arrived at Edinburgh Zoo amid worldwide interest on December 4 last year after a 5,000-mile flight from China.
Since then, more than half a million visitors have been to see the animals, who are set to stay in the country for another nine years.
Iain Valentine, director of research and conservation at the zoo, said much has been learnt about the species over the last 12 months.
"Looking back on this, our first giant panda year, it has been a great success," he said.
"Although we are a world leader in the science and art of animal husbandry, we at the Royal Zoological Society of Scotland (RZSS) have learnt an enormous amount about Tian Tian and Yang Guang, but also about giant pandas as an entire species. This knowledge is vital to the global conservation panda effort.
"The core focus of RZSS as a charity is conservation, research and education. Over the period of the last 12 months we have achieved a great deal in these crucial areas, of which we are extremely proud. Over half a million people have visited the pandas from Scotland, the UK and further afield, that translates to over half a million people that we have educated about pandas through our guided tours around the panda exhibit. That's a phenomenal reach and impact and one that I hope will continue long into the future."
Zoo bosses said the arrival of female panda Tian Tian, whose name means "sweetie", and male Yang Guang, meaning "sunshine", was the result of five years of political and diplomatic negotiations.
The pandas, who have a limited breeding season, did not manage to mate this year.
The zoo said it believes Yang Guang came into season two weeks after the female and that next year keepers will adhere to a strict "lights out" policy to try to synchronise the animals' breeding cycles.